Stan Braude

Stan Braude

​Professor of the Practice of Biology
PhD University of Michigan
Animal Behavior Society Distinguished Teacher 2011
National College Biology Teacher of the Year 2004

contact info:

mailing address:

  • Washington University
    CB 1137
    One Brookings Drive
    St. Louis, MO 63130-4899

​Stan Braude is a professor of the practice of biology. A current project focuses on endangered Hine's emerald dragonfly populations in Missouri.

Stan Braude’s primary research focuses on the evolution of social behavior and senescence in naked mole-rats in east Africa.  He has also worked on  Mecopteran and Panorpa scorpionflies, Patagonian Tuco-tucos, rhinoceros inbreeding, Hines Emerald dragonfly demography, domestication of dogs, and ultrasound diagnostics in Olms.  In 2019 he begins work on giant pouch rats in Tanzania.  Here at Wash U, Braude teaches Human Anatomy and Physiology, Advanced Wilderness Medicine, Missouri’s Natural Heritage, the Biology of Dog Breeds, and in 2020 he will begin offering the Biology of Trees.

recent courses

First-Year Opportunity: The Biology of Dog Breeds

This freshman seminar uses the topic of dog behavior and genetics to teach fundamental scientific tools and to engage students in contributing to the building of an online public resource that summarizes the scientific literature on breeds. Our first task is learning to read and dissect primary scientific literature. We parse out the difference between scientific questions, hypotheses, and predictions through a guided case-study exercise. We then apply the experience to outlining primary research articles, identifying the key components of the author's arguments and summarizing the results and implications. The second half of the semester is spent searching the scientific literature, sorting information into the new dog breed resource, and presenting results to peers around the seminar table.

    AMPERSAND: Missouri's Natural Heritage, part 1

    Missouri's Natural Heritage is for freshmen who want to get outdoors and learn about their home for the next four years. The first semester of the sequence will focus on Missouri geology, climate, archaeology, and native megafauna. This will provide a foundation on which to examine the ecology, restoration, and management of our diverse habitats (prairie, forest, glade, and stream) and the biology of our diverse plant and animal wildlife (arthropods, mollusks, fish, salamanders, lizards, birds, and mammals) in the second semester. We will also introduce basic concepts in biodiversity and resource management with case studies from here in Missouri. In addition to weekly lecture and discussion, students in this class will visit sites across the state during a number of Friday afternoon field trips and weekend camping trips. Camping gear, transportation and meals for all field trips are covered by the lab fee.

      Introduction to Anatomy and Physiology I

      This is the first of a two-semester sequence that examines all major organ systems in the human/mammalian body. The lab is an integral part of the course. The emphasis is on understanding normal function and processes at the gross, cellular, and molecular levels as well as some discussion of pathology and disease. The first semester covers basic principles of cellular physiology, histology, digestion, bone, muscle, and nervous systems. Optional weekly discussion and review sections are also offered during which case studies are discussed as a means of applying and reviewing lecture material.

        Advanced General Physiology

        This graduate-level course will examine physiology in a cohesive evolutionary context. For each system we will first review the general anatomy and physiology, turning then to the molecular basis of function and dysfunction. Each class provides an overview of the basic physiology of one of the major organ systems including: neural, respiratory, cardiovascular, endocrine, digestive, urogenital and immune, followed by a discussion of recent discoveries presented in primary scientific literature and current reviews.

          Selected Publications


          Braude, S. and B. Low. 2010. An Introduction to Methods and Models in Ecology, Evolution, and Conservation Biology. Princeton University Press, Princeton, NJ.

          Braude, S., A. Miceli, and D. Goran. 2012. Case Studies for Understanding the Human Body, Second Edition.  Jones and Bartlett, Sudbury, MA.


          Ingram, C., Troendle, N., Gill, C., Braude, S., and Honeycutt, R., 2015. Challenging the inbreeding hypothesis in a eusocial mammal: population genetics of the naked mole-rat, Heterocephalus glaber.  Molecular Ecology 24, 4848–4865.

          Braude, S. and A. Beck.  2013.  Complete Blood Counts with Differential: more accurate reference ranges based on circadian leukocyte trafficking.  Journal of Clinical Pathology 66:109-110.

          Braude, S. and J. Gladman.  2013.  Out of Asia:  An Allopatric Model for the Evolution of the Domestic Dog. ISRN Zoology 2013:1-7

          Braude, S. 2013.  Chapter 12: The adaptive significance of humor.  In:  Foundations of human social evolution:  Contributions of Richard D. Alexander.  Crespi, B. and Summers, K.,  eds.  Oxford University Press, New York.

          Braude, S. and R. Templeton. 2009 Understanding the Multiple Meanings of Inbreeding and Effective Size for Management of Rhinoceros Populations. African Journal of Ecology 47: 546-555.

          Goran, D. and Braude, S. 2007. Social and Cooperative Learning in the Solving of Case Histories. American Biology Teacher, 69 (2): 80-84.

          Braude, S. 2007. The Multiple Meanings of Inbreeding: The Key to Understanding the Social and Genetic Structure of Subterranean Rodent Populations. In Beagal, S., Burda, H., Schleich, C., eds. Subterranean Rodents: News from Underground. Springer Verlag. pp 331-340.

          Braude, S. and L. Corey. 2006. The Confidence Game: A simple activity with dice teaches students the importance of replication and sample size in data collection. Science and Children 44(1):40-44.

          Braude, S., Ciszek, D., Berg, N., and Shefferley, N., 2001. The Ontogeny and Distribution of Countershading in Colonies of the Naked mole-rat, Heterocephalus glaber. Journal of Zoology London 253: 351-357.

          Braude, S. 2000. Dispersal and new colony formation in wild naked mole-rats: evidence against inbreeding as the system of mating. Behavioral Ecology 11:7-12.

          Braude, S., Tang-Martinez, Z., and Taylor, G. 1999 Stress, Testosterone, and the Immunoredistribution Hypothesis. Behavioral Ecology. 10(3): 345-350.

          Sherman, P., Braude, S., and Jarvis, JUM, 1999. Litter sizes and mammary numbers of naked mole-rats: breaking the one-half rule. Journal of Mammalogy 80(3):720-733

          Braude, S. 1998. The predictive power of evolutionary biology and the discovery of eusociality in the naked mole-rat. NCSE Reports, 17(4): 12-15.

          Lacey, E., Braude, S., and Weiczorek, J. 1997. Burrow sharing by colonial tuco-tucos (Ctenomys sociabilis). Journal of Mammalogy 78(2): 556-562.